Lactuca glandulifera

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Lactuca glandulifera

http://media.bgbm.org/erez/erez?src=EditWP6/photos/Lactuca_glandulifera_A_01.jpg

Description

Perennial scrambling or scandent herb, 0.3-2 m high; stems to 3 m long, distally leafy, viscid with dense glands but glabrescent, reddish when young, with abundant white latex; older stems with characteristic papery decorticating bark. Leaves variously lobed, 2-12 cm long, 0.8-5.5 cm wide, the very base sagittate-semi- amplexicaul, continuing as a winged pseudopetiole widening into a pair (rarely two pairs) of retrorse triangular lobes 0.2-3 cm long, the terminal lobe triangular to ovate and 1-7 cm long, margins sinuate to dentate to almost lobed, apex acute or rounded, densely glandular-setose, especially on midrib and margins, but glabrescent. Capitula many, in terminal leafy panicles to 30 cm long with ± dichotomous branching; involucre 8-15 mm long, cylindrical at anthesis becoming narrowly conical in fruit; phyllaries dark; green tinged with red or purple, linear or elliptic, 2-15 mm long, obtuse or rounded, sparsely setose or glabrous, with a tiny distal hair-tuft. Florets 4-5 per capitulum; Corolla pale yellow or less often white, sometimes with purple tinge, tube cylindric, 4-6 mm long, ligule 5-9 mm long, 1.5-2.5 mm wide. Achenes pale brown, ellipsoid and (slightly) compressed, 5.3-6 mm long including a 1-2 mm long beak, 3-6-ribbed on each side, glabrous or puberulous; pappus white, 5.5-8 mm long.

slightly modified (for floret number) from: Jeffrey, C. & Beentje, H. J. 2000: Cichorieae. – Pp. 63-108 in: Beentje, H. J. & Smith, S. A. L. (ed.), Flora of Tropical East Africa. Compositae (Part 1). – Kew.

Systematics

Lactuca elgonensis has been included in L. glandulifera by Beentje & Jeffrey (2000) while according to the view maintained here, it should be treated as a separate species, for details see under L. elgonensis. A

Distribution

Africa: Burundi (BurundinativeB,C,D); Cameroon (CameroonnativeA,B,D,E,F,G,H,I,J); Ethiopia (EthiopianativeA,B,C,D,F,H,I,K); Gulf of Guinea Is. (BiokonativeA,B,F,I,L,M); Ivory Coast (Ivory CoastnativeB,D); Kenya (KenyanativeA,B,F,I,N); Liberia (LiberianativeB,D); Malawi (MalawinativeA,B,C,D,H,I,K,O); Mozambique (MozambiquenativeA,B,C,D,H,I,K); Nigeria (NigerianativeA,F,I,P); Rwanda (RwandanativeA,B,C,D,I,Q); Sierra Leone (Sierra LeonenativeA,B,D,F,H,I); Sudan (SudannativeD,H,R); Tanzania (TanzanianativeA,B,F,H,I); Uganda (UgandanativeA,B,F,I,S); Zaire (ZairenativeA,B,C,D,F,I,T,U)

Common names

Kinyarwanda (Central and East Africa): RuvunangeB, UruvunankaB; Mashi (Nigeria) (Central and East Africa): LuvunangaB, VunangaB; Nilo-Saharan (Other) (Central and East Africa): DyedyembidyasiB; Oku (Cameroon): IyenG; Rundi (Central and East Africa): IbitekamatekeB

Credits

Zhang J.-W. 2017: Images (3 made by Yan-Bo Li and Dao Teng added)

Bibliography

A. Beentje H. J. & Jeffrey C. - Cichorieae in Polhill R.M. (ed.), Flora of Tropical East Africa. Compositae 1. 2000
B. Lawalrée A., Dethier D. & Gilissen E., Flore d'Afrique Centrale (Zaire-Rwanda-Burundi). Spermatophytes, Compositae I. 1986
C. Pope G. V., Flora Zambesiaca 6(1). 1992
D. Tadesse M. - 16. Scolymus - 29. Tragopogon in Hedberg I. & Edwards S., Flora of Ethiopia and Eritrea. 2004
E. Cheek M., Pollard B. J., Darbyshire I., OnanaJ.-M. and Wild C., The Plants of Kupe, Mwanenguba and the Bakossi Mountains, Cameroon. 2004
F. Adams C. D. - Tribe XIII Cichorieae in Hutchinson J. & Dalziel J. M. (ed.), Flora of West Tropical Africa, ed. 2, 2. 292-297
G. Beentje H. J., Onana J.-M., Pollard B. J. & Hind D. J. N. - Compositae in Cheek M., Onana J.-M. & Pollard B. J. (ed.), The plants of Mount Oku and the Ijim Ridge, Cameroon. A conversation checklist. 2000
H. Friis I. & Vollesen K., Flora of the Sudan-Uganda border area east of the Nile in Biol. Skr. 51(1). 1998
I. Jeffrey C., Notes on Compositae: 1. The Cichorieae In East Tropical Africa in Kew Bull. 18 (3). 1966
J. Oliver D. & Hiern W. P. - Compositae in Oliver D., Flora of Tropical Africa 3. 1877
K. Burrows J. E. & Willis C. K., Plants of Nyika Plateau. An account of the vegetation of the Nyika National Parks of Malawi and Zambia in SABONET Report 31. 2005
L. Aedo C. & al., Bases documentales para la flora de Guinea ecuatorial. 1999 (as Lactuca glandulifera)
M. Velayos M. & al., Preliminary checklist of vascular plants of Bioko Island in Bot. Complut. 37. 2013
N. Agnew A. D. Q. & Agnew S., Upland Kenya wild flowers. A flora of the ferns and herbaceous flowering plants of Upland Kenya, ed. 2. 1994
O. Adams C. D. - Tribe XIII Cichorieae in Hutchinson J. & Dalziel J. M. (ed.), Flora of West Tropical Africa, ed. 2, 2. 292-297 (as Lactuca glandulifera f. calva R. E. Fr.)
P. Adams C. D. - Tribe XIII Cichorieae in Hutchinson J. & Dalziel J. M. (ed.), Flora of West Tropical Africa, ed. 2, 2. 292-297 (as Lactuca glandulifera var. calva (R. E. Fr.) Robyns)
Q. Lawalrée A. - 56. Tolpis - 61. Crepis in Troupin G., Flore du Rwanda. Spermatophytes 3. 1985
R. Darbyshire I. & al., The plants of Sudan and South Sudan. An annotated checklist. 2015
S. Jeffrey C., Notes on Compositae: 1. The Cichorieae In East Tropical Africa in Kew Bull. 18 (3). 1966 (as Lactuca elgonensis Stebbins)
T. Robyns W., Flore des Spermatophytes du Parc National Albert. Sympétales 2. 1947 (as Lactuca glandulifera var. calva (R. E. Fr.) Robyns)
U. Robyns W., Flore des Spermatophytes du Parc National Albert. Sympétales 2. 1947 (as Lactuca wildemaniana Stebbins)