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The molecular analyses by Gemeinholzer & al. (in Kilian & al. 2009) revealed a clade comprising the genera Chondrilla, Phitosia and Willemetia, which has been recognised as a separate subtribe Chondrillinae by Kilian & al. (2009). Whereas the sister-group relationship of Chondrilla and Willemetia received strong statistical support, there was no statistical support for the clade also including Phitosia, a monospecific genus recently established for a species removed from Crepis for cytological and carpological reasons (Kamari & Greuter 2000).

Chondrilla and Willemetia have been hitherto treated as members of subtribe Crepidinae (Bremer 1994; Lack 2006). Also in the ITS phylogeny by Gemeinholzer & al. (in Kilian & al. 2009) the Chondrillinae and Crepidinae clades together formed a monophyletic group with full bootstrap and posterior probability support. However, since the sister-group relationsship of Crepidinae and Chondrillinae has been proven less stable when different molecular datasets (matK and nrITS) were combined (Gemeinholzer & al. unpubl.), Kilian & al. (2009) refrained from treating the Chondrilla alliance as a part of the subtribe Crepidinae.


Bremer K. 1994: Asteraceae. Cladistics and classification. – Portland: Timber.

Kilian N., Gemeinholzer B. & Lack H. W. 2009: Tribe Cichorieae. – In: Funk V. A., Susanna A., Stuessy T. & Bayer R. (ed.), Systematics, evolution, and biogeography of the Compositae. – Vienna: IAPT.

Kamari G. & Greuter W. 2000: Phitosia, a new genus for Crepis crocifolia (Compositae, Cichorieae), a local endemic of Mount Taigetos (Greece). – Bot. Chron. (Patras) 13: 11-36.

Lack H. W. 2006: Tribe Cichorieae Lam. & DC. – Pp. 180-199 in: Kadereit J. W. & Jeffrey C. (ed.), The families and genera of vascular plants 8. – Berlin: Springer.


Africa: Algerianative; Libyanative; Morocconative; Tunisianative Asia-Temperate: Afghanistannative; Altaynative; Cyprusnative; East Aegean Is.native; Irannative; Iraqnative; Kazakhstannative; Kirgizistannative; Lebanon-Syria (Lebanonnative, Syrianative); Mongolianative; North Caucasusnative (Dagestannative, Krasnodarnative, Stavropolnative); Palestine (Israelnative, Jordannative); Saudi Arabianative; Tadzhikistannative; Tibetnative; Transcaucasus (Abkhaziyanative, Adzhariyanative, Armenianative, Azerbaijannative, Georgianative, Nakhichevannative); Turkeynative; Turkmenistannative; Uzbekistannative; West Siberianative; Xinjiangnative Asia-Tropical: Pakistannative; West Himalaya (Jammu-Kashmirnative) Australasia: New South Wales (Australian Capital Territoryintroduced, New South Walesintroduced); New Zealand Northintroduced; Northern Territoryintroduced; Queensland (Queenslandintroduced); South Australiaintroduced; Tasmaniaintroduced; Victoriaintroduced; Western Australia (Western Australiaintroduced) Europe: Albanianative; Austrianative (Austrianative, Liechtensteinnative); Balearesnative; Baltic States (Kaliningradnative, Latviaintroduced, Lithuaniaintroduced: adventitious (casual)); Belarusnative; Belgium (Belgiumnative, Luxembourgnative); Bulgarianative; Central European Russianative; Corsenative; Czechoslovakia (Czech Republicnative, Slovakianative); East European Russianative; Francenative; Germanynative; Greecenative; Hungarynative; Italynative; Kritinative; Krymnative; Netherlandsnative: formerly native; Polandnative; Portugalnative; Romanianative; Sardegnanative; Sicilynative (Sicilynative); South European Russianative; Spainnative (Andorranative, Gibraltarnative, Spainnative); Switzerlandnative; Turkey-in-Europenative; Ukraine (Moldovanative, Ukrainenative); Yugoslavia (Bosnia-Herzegovinanative, Croatianative, Macedonianative, Montenegronative, Serbianative, Slovenianative) Northern America: British Columbiaintroduced; Californiaintroduced; Delawareintroduced; District of Columbiaintroduced; Georgia, U.S.A.introduced; Idahointroduced; Illinoisintroduced; Marylandintroduced; Michiganintroduced; Montanaintroduced; New Jerseyintroduced; Ontariointroduced; Oregonintroduced; Pennsylvaniaintroduced; Virginiaintroduced; Washingtonintroduced; West Virginiaintroduced Southern America: Argentina Northeast (Buenos Airesintroduced)